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Hirotoshi Tsuda, Hisato Kunitake, Mai Yamasaki, Haruki Komatsu, and Katsunori Yoshioka

( Lyrene, 2011 ). In our previous study, we have succeeded to obtain several tetraploid shashanbo plants by colchicine treatment (H. Tsuda, unpublished data). The objectives of the present study were as follows: 1) to produce intersectional hybrids from the

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Hirotoshi Tsuda, Hisato Kunitake, Yo Aoki, Akiko Oyama, Takuya Tetsumura, Haruki Komatsu, and Katsunori Yoshioka

to the section Cyanococcus ( Karizumi, 1979 ; Kunitake et al., 2006 ; Luby et al., 1991 ). Therefore, we produced intersectional hybrids between colchicine-induced tetraploid shashanbo and the tetraploid highbush blueberry ‘Spartan’ ( V

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Paul M. Lyrene

combinations in Vaccinium is very large. Relatively few attempted combinations have been reported, and no hybrid seedlings were produced in some reports. Intersectional hybrids obtained by crossing diploid Vaccinium species have usually been sterile or

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Paul M. Lyrene

the tetraploids being the result of unreduced gametes from the diploid parent. A few intersectional Vaccinium hybrids have been discovered or created. The most widely studied natural intersectional hybrid {[ V. intermedium = V. myrtillus (section

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P.M. Lyrene

Diploid plants in Vaccinium section Cyanococcus, including plants of V. darrowi Camp, V. atrococcum. Heller (diploid V. corymbosum L.), a V. atrococcum × V. darrowi F1 hybrid, and a V. atrococcum × V. elliottii Chapmn. F1 hybrid, were hand pollinated in a greenhouse with pollen from diploid V. arboreum Marsh. (Section Batodendron). The resulting seeds were germinated and the seedlings were transplanted to a high-density field nursery. Forty of these F1 intersectional hybrids were selected after 2 1/2 years and transplanted to a 1.5 × 4-m spacing. Most of these plants were vigorous and flowered heavily in subsequent years, but only a small percentage of the flowers produced fruit. In 1990, however, >4000 berries were harvested from the 35 surviving plants. Open-pollinated seed from a much smaller number of berries was planted in Dec. 1987; these seeds produced ≈200 seedlings, some of which had moderate to high fruit set in a field nursery in 1989. Six of these seedlings, which were selected for high vigor, high fruit set, and characteristics intermediate between section Cyanococcus and section Batodendron, had fruit set ranging from 19.4% to 92.7% when pollinated with pollen from tetraploid V. corymbosum cultivars. One of the six seedlings was highly self-fruitful, and some intercrosses among the six seedlings produced much viable seed. Large-scale introgression of V. arboreum genes into tetraploid highbush cultivars likely will be possible by the methods used in this study.

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Paul M. Lyrene

× V. arboreum crosses made later using other tetraploid V. arboreum plants. These crosses gave far fewer hybrids per pollinated flower than analogous intersectional hybrids made at the diploid level ( V. darrowii section Cyanococcus × V

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Bruce L. Dunn and Jon T. Lindstrom

color for T. arizonicum × T. lanatum hybrids were purple, and thus resemble the purple-blue flowers of T. lanatum . Table 1. Morphological comparisons among T. arizonicum , the intersectional hybrid T . arizonicum × T. lanatum, T

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Mark K. Ehlenfeldt and James L. Luteyn

remains to be understood about intersectional hybrids in Vaccinium ; however, given these results and the results of several other studies ( Ehlenfeldt and Polashock, 2014 ; Lyrene, 2011 , 2016 ; Morozov, 2007 ), intersectional crossing barriers appear

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Barbara Gilmore, Nahla Bassil, April Nyberg, Brian Knaus, Don Smith, Danny L. Barney, and Kim Hummer

types. Intersectional hybrids are crosses between the two groups. Intersectionals (Itohs) have a similar growth pattern to herbaceous peonies, so they are able to withstand temperatures that would kill tree peonies, but their foliage and the flowers have

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Paul M. Lyrene

hybrids was given ( Lyrene, 2016 ). The purpose of this article is to describe the results of backcrossing these F 1 intersectional hybrids to the two parent taxa. Materials and Methods Evaluation of hybrid seedlings in high-density field nursery. Seeds