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Chad E. Finn, Bernadine C. Strik, Brian M. Yorgey, Mary E. Peterson, Patrick A. Jones, Jungmin Lee, Nahla V. Bassil, and Robert R. Martin

-erect blackberries like ‘Triple Crown’ ( Galletta et al., 1998b ). ‘Ouachita’ and ORUS 2867-4 trace their thornlessness to ‘Merton Thornless’ ( Crane, 1943 ; Jennings, 1986 ). ‘Ouachita’ and ‘Triple Crown’ are tetraploid (2 n = 4 x = 28), while the grandparent

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Chad E. Finn, Bernadine C. Strik, Brian M. Yorgey, Mary E. Peterson, Patrick A. Jones, Gil Buller, Jungmin Lee, Nahla V. Bassil, and Robert R. Martin

at the hexaploid or higher ploidy levels produced fewer seeds than typical and no viable seedlings in the field. Similar to other erect and semierect thornless cultivars, ‘Galaxy’ has the ‘Merton Thornless’ source of thornlessness (botanically

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Brent Black, James Frisby, Kimberly Lewers, Fumiomi Takeda, and Chad Finn

twofold difference in GDH to bloom among genotypes, indicating significant genetic variation ( Fig. 3 ). Among cultivars in the collection, GDH to bloom varied from 9,200 GDH for ‘Chickasaw’ to 18,900 GDH for ‘Merton Thornless’ based on a linear model with

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Jason D. Zurn, Katie A. Carter, Melinda H. Yin, Margaret Worthington, John R. Clark, Chad E. Finn, and Nahla Bassil

. hochstetterorum ( Lopes et al., 2006 ), and eight from ‘Marion’ blackberry ( Castillo et al., 2010 ). Lewers et al. (2008) generated expressed sequence tags (ESTs) from ‘Merton Thornless’ blackberry and predicted the presence of 940 polymorphic SSRs in the

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Nina R.F. Castillo, Barbara M. Reed, Julie Graham, Felicidad Fernández-Fernández, and Nahla Victor Bassil

reliable means for cultivar identification and to assess genetic relatedness and diversity in these collections. Microsatellite markers were only recently developed from an expressed sequence tag library of ‘Merton Thornless’ ( Lewers et al., 2008 ). A

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Chad E. Finn, Bernadine C. Strik, Brian M. Yorgey, Mary E. Peterson, Jungmin Lee, Robert R. Martin, and Harvey K. Hall

in breeding of the various sources of thornlessness have been well reviewed ( Clark et al., 2007 ; Finn and Clark, 2012 ; Hall and Stephens, 1999 ). To briefly compare the sources of thornless, the ‘Merton Thornless’ source, derived from ‘John Innes

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Chad E. Finn, Bernadine C. Strik, Brian M. Yorgey, Mary E. Peterson, Patrick A. Jones, Gil Buller, Sedat Serçe, Jungmin Lee, Nahla V. Bassil, and Robert R. Martin

et al., 1998a ) that are standard cultivars in the industry. As with the other erect and semierect thornless cultivars, ‘Eclipse’ has the ‘Merton Thornless’ source of thornlessness (botanically “spineless” but commonly referred to as “thornless” in

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Alisha L. Ruple, John R. Clark, and M. Elena Garcia

degeneration and reduces flower set in sweet cherry Sci. Hort. 119 455 457 Hellman, E.W. Clark, J.R. 1984 Blackberry pollen sterility associated with ‘Merton Thornless’ germplasm Fruit Var. J. 38 55 57

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Ksenija Gasic, John E. Preece, and David Karp

to Chester Thornless; stores very well. Plant: growth habit semi-erect to very erect; crown forming; thornless with the Merton Thornless source of thornlessness; vigorous; yield moderate, 6-7.2 kg/plant; less susceptible to UV and heat damage than

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Ksenija Gasic, John E. Preece, and David Karp

; fruit store very well with low levels of drupelet reversion; ripens after Eclipse and Galaxy, before Triple Crown and well before Chester Thornless. Plant: vigorous; growth habit semi-erect; crown forming; thornless with the Merton Thornless source of