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Jack B. Fisher, Anders Lindström, and Thomas E. Marler

scale ( Aulacaspis yasumatsui Takagi) is among the greatest of threats ( IUCN Cycad Specialist Group, 2006 ). It has thrust natural populations of many species into a vulnerable position and devastated the international horticultural trade built on

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Thomas E. Marler and Nirmala Dongol

the armoured scale Aulacaspis yasumatsui J. Trop. Ecol. 28 233 242 Marler, T.E. Lindström, A.J. 2014 Free sugar profile in cycads Front. Plant Sci. 5 526 Marler, T.E. Lindström, A.J. 2015 Carbohydrates, pollinators, and cycads Commun. Integr. Biol. 8

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Thomas E. Marler, Vivian Lee, and Christopher A. Shaw

the most abundant forest species to being “red-listed” as endangered by the IUCN ( Marler et al., 2006 a) because of the assault by the invasive scale, Aulacaspis yasumatsui Takagi ( Haynes, 2006 ; Marler and Muniappan, 2006 ). The urgency of

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Thomas E. Marler, Anders Lindström, and Jack B. Fisher

for use on six Cycas species to evaluate efficacy for control of Aulacaspis yasumatsui Takagi and resolve how the pachycaulous stem would respond to the treatment. We determined that the stem's succulent nature was highly susceptible to secondary

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Trenton Hamada, Irene Terry, Robert Roemer, and Thomas E. Marler

damage by invasive cycad specific pests, the scale Aulacaspis yasumatsui Takagi (Hemiptera: Diaspididae) ( Marler and Muniappan, 2006 ), and the cycad blue butterfly, Chilades pandava Horsfield (Lepidoptera: Lycaenidae) ( Moore et al., 2005 ). Surveys

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Benjamin E. Deloso, Anders J. Lindström, Frank A. Camacho, and Thomas E. Marler

micronesica following Aulacaspis yasumatsui infestation HortScience 53 1125 1128 Marler, T.E. Lindström, A. Fisher, J.B. 2010 Stem tissue dimensions correlate with ease of horticultural management for six Cycas species HortScience 45 1293 1296 Marler, T

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Thomas E. Marler and Christopher A. Shaw

secondary to that of the exotic Cycas revoluta Thunb. However, both species have been decimated from the horticultural scene as a result of the 2003 invasion of the cycad-specific scale Aulacaspis yasumatsui Takagi. This arthropod invasion positioned C