Received for publication May 15, 1980. Scientific Journal Series Paper Number 11222 of the Minnesota Agricultural Experiment Station. This study was supported by grant USDI No. PX-6000-7-0924 from the United States National Park Service. We gratefully acknowledge donations of seeds from Dr. Joseph Arditti, University of California, Irvine, Calif.; Fredrick W. Case, Saginaw, Mich.; Joyce Cloughly, Basking Ridge, N.J.; Ruth and Hugh Cocker, Rochester, Minn.; Betty Mech, Minneapolis, Minn.; Sandy Murphy, Edina, Minn.; Jan Norman, Golden Valley, Minn.; University of Minnesota Landscape Arboretum, Chaska, Minn.; and the University of Toronto, Toronto, Ontario (Canada).
Present address: Missouri Botanical Garden, Botany Department, P.O. Box 299, St. Louis, MO 63166.
Present address: Department of Horticulture, University of Maryland, College Park, MD 20742.
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