Received for publication Feb. 16, 1982. Paper number 12,074 of the Scientific Journal Series, University of Minnesota Agricultural Experiment Station. Research was supported by grant USDI No. PX-6000-7-0924 from the U.S. National Park Service. We gratefully acknowledge J. Arditti, Irvine, Calif., who donated seeds of Epipactis gigantea Douglas ex. Hooker.
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